[1] Bull JJ.Temperature-sensitive periods of sex determination in alizard:comparisons with turtles and crocodilians[J] . Journal of Experimental Zoology, 1987, 241:143-149. [2] Grossen C, Neuenschwander S, Perrin N.Temperature-dependent turnovers in sex-determination mechanisms:a quantitative model[J] . Journal of Evolutionary Biology, 2011, 65(1):64-78. [3] Craves, JAM.Weird animal genomes and the evolution of vertebrate sex and sex chromosomes[J] . The Annual Review of Genetics, 2008, 42:565-586. [4] Valenzuela N, Adams DC, Janzen FJ.Pattern does not equal process:exactly when is sex environmentally determined[J] ? Amrican Naturalist, 2003, 161:676-683. [5] Guler Y, Short S, Kile P, et al.Integrating field and laboratory evidence for environmental sex determination in the amphipod, Echinogammarus marinus[J] . Marine Biology, 2012, 159:2885-2890. [6] Kobayashi Y, Nagahama Y, Nakamura M.Diversity and plasticity of sex determination and differentiation in fishes[J] . Sexual Development, 2013, 7:115-125. [7] Becak W, Becak ML, Nazareth HR.Karyotypic studies of two species of South American snakes(Boa constrictor amarali and Bothrops jararaca)[J] . Cytogenetics, 1962, 1(6):305-313. [8] Bull JJ.Sex determination in reptiles[J] . Quarterly Review of Biology, 1980, 55:3-21. [9] Robert KA, Thompson MB.Sex determination-Viviparous lizard selects sex of embryos[J] . Nature, 2001, 412(6848):698-699. [10] Bull JJ, Bulmer MG.The evolution of XY females in mammals[J] . Heredity, 1981, 47(3):347-365. [11] Ezaz T, Valenzuela N, Grutzner F, et al.An XX/XY sex microchro-mosome system in a freshwater turtle, Chelodina longicollis(Testudines:Chelidae)with genetic sex determination[J] . Chromosome Research, 2006, 14(2):139-150. [12] Wapstra E, Warner DA.Sex allocation and sex determination in squamate reptiles[J] . Sexual Development, 2010, 4(1-2):110-118. [13] 史海涛. 中国的龟类. 生物学通报[J]. 2004, 39(5):13-17. [14] 周婷, 李丕鹏. 中国龟鳖物种多样性及濒危现状[J]. 四川动物, 2007, 26(2):463-467. [15] 刘凌云. 普通动物学[M]. 第3版. 北京:高等教育出版社, 1997, 50-57. [16] Mccord WP, Iverson JB, Spinks PQ.A new genus of geoemydid turtle from Asia[J] . Hamadryad, 2000, 25(2):20-24. [17] Zhou X, Zhu Y.Sex-specifc growth characteristics of Chinese sof-shelled turtle, Pelodiscus sinensis[J] . Aquaculture, 2011, 5:11-13. [18] Kawagoshi T, Uno Y, Matsubara K, et al.The ZW micro-sex chromosomes of the Chinese soft-shelled turtle(Pelodiscus sinensis, Trionychidae, Testudines)have the same origin as chicken chromosome 15[J] . Cytogenet and Genome Research, 2009, 125(4):125-131. [19] Mu Y, Zhao B, Tang WQ, et al.Temperature-dependent sex determination ruled out in the Chinese soft-shelled turtle(Pelodiscus sinensis)via molecular cytogenetics and incubation experiments across populations[J] . Sexual Development, 2015, 9(2):111-117. [20] 聂刘旺, 郭超文, 汪鸣, 王千. 中华鳖的性别决定机制[J]. 应用与环境生物学报, 2001, 7(3):258-261. [21] 朱道玉, 孙西寨. 中华鳖的性别决定[J]. 动物学杂志, 2000, 35(6):37-39. [22] 高丽丽. 中华鳖Foxl2基因克隆与表达及温度和外源激素对其表达的影响[D]. 重庆:西南大学, 2016. [23] 杨振才, 牛翠娟, 孙儒泳. 中华鳖生物学研究进展[J]. 动物学杂志, 1999, 34(6):42-45. [24] 康辰香, 闫志民. 龟鳖. 北京:中国中医药出版社, 2000:151-152. [25] Liang L, Sun BJ, Ma L, et al.Oxygen-dependent heat tolerance and developmental plasticity in turtleembryos[J] . Journal of Comparative Physiology B-Biochemical Systemic and Environmental Physiology, 2015, 185(2):257-263. [26] Charnier M.Action of Temperature on Sex-Ratio in Agama Agamas Embryo(Agamidae Lacertilien)[J] . Comptes Rendus Des Seances De LA Societe De Bioligie Et De Ses Filiales, 1966, 160(3):620-622. [27] Pezaro N, Thompson MB, Doody JS.Seasonal sex ratios and the evolution of temperature-dependent sex determination in oviparous lizards[J] . Evolutionary Ecology, 2016, 30(3):551-565. [28] Wapstra E, Olsson M, Shine R, et al.Maternal basking behaviour determines offspring sex in a viviparous reptile[J] . Proceedings of The Royal Society B-Biological Sciences, 2004, 7(271):230-232. [29] Langkilde T, Shine R.Different optimal offspring sizes for sons versus daughters may favor the evolution of temperature-dependent sex determination in viviparous lizards[J] . Evolution, 2005, 59(10):2275-2280. [30] Ewert MA, Jackson DR, Nelson CE.Patterns of temperature-dependent sex determination in turtles[J] . Journal of Experimental Zoology, 1994, 270(270):3-15. [31] Inamdar Doddamani LS, Vani V, Seshagiri PB.A tropical oviparous lizard, Calotes versicolor, exhibiting a potentially novel FMFM pattern of temperature-dependent sex determination[J] . Journal of Experimental Zoology Part A-Ecological Genetics and Physiology, 2012, 317(1):32-46. [32] Bull JJ.Temperature-sensitive with turtle and crocodilians[J] . The Journal of Experimental Zoology, 1987, 241:143-145. [33] Viets BE, Tousignant A, Ewert MA, et al.Temperature-dependent sex determination in the leopard gecko, Eublepharis macularius[J] . Journal of Experimental Zoology, 1993, 265(6):679-683. [34] Booth DT.Influence of incubation temperature on hatchling phenotype in reptiles[J] . Physiological and Biochemical Zoology, 2006, 79(2):274-281. [35] Rhen T, Fagerlie R, Schroeder A, et al.Molecular and morphological differentiation of testes and ovaries in relation to the thermosensitive period of gonad development in the snapping turtle, Chelydra serpentina[J] . Differentiation, 2015, 89(1-2):31-41. [36] Matsumoto Y, Crews D.Molecular mechanisms of temperature-dependent sex determination in the context of ecological developmental biology[J] . Molecular and Cellular Endocrinology, 2012, 354(1-2):103-110. [37] Matsumoto Y, Yatsu R, Taylor C, et al.Changes in gonadal gene network by exogenous ligands in temperature-dependent sex determination[J] . Journal of Molecular Endocrinology, 2013, 50(3):389-400. [38] Braña F, Ji X.Influence of incubation temperature on morphology, locomotor performance, and early growth of hatchling wall lizards(Podarcis muralis)[J] . Journal of Experimental Zoology, 2000, 286(4):422-433. [39] Navarro-Martín L, Viñas J, Ribas L, et al.DNA methylation of the gonadal aromatase(cyp19a)promoter is involved in temperature-dependent sex ratio shifts in the European sea bass[J] . PLoS Genetics, 2011, 7(12):e1002447. [40] Warner DA, Mitchell TS, Bodensteiner BL, et al.The effect of hormone manipulations on sex ratios varies with environmental conditions in a turtle with temperature-dependent sex determination[J] . Journal of Experimental Zoology Part A- Ecological Genetics and Physiology, 2017, 327(4):172-181. [41] Sun BJ, Li T, Mu Y, et al.Thyroid hormone modulates offspring sex ratio in a turtle with temperature-dependent sex determination[J] . Proceedings of The Royal Society B-Biological Sciences, 2016, 283(1841):20161206. [42] Conover DO.Adaptive significance of temperature-dependent sex determination in a fish[J] . American Naturalist, 1984, 123, 297-313. [43] Warner DA, Shine R.The adaptive significance of temperature-dependent sex determination in a reptile[J] . Nature, 2008, 451(7178):566-568. [44] Freedberg S, Taylor DR.Sex ratio variance and the maintenance of environmental sex determination[J] . Journal of Evolutionary Biology, 2007, 20(1):213-220. [45] Freedberg S, Wade MJ.Male combat favours female-biased sex ratios under environmental sex determination[J] . Animal Behaviour, 2004, 67:177-181. [46] Mitchell TS, Maciel JA, Janzen FJ.Does sex-ratio selection influence nest-site choice in a reptile with temperature-dependent sex determination[J] ? Proceedings of The Royal Society B-Biological Sciences, 2013, 280(1772):20132460. [47] Wapstra E, Uller T, While GM, et al.Giving offspring a head start in life:field and experimental evidence for selection on maternal basking behaviour in lizards[J] . Journal of Evolutionary Biology, 2010, 23(3):651-657. [48] Radder RS, Pike DA, Quinn AE, et al.Offspring sex in a lizard depends on egg size[J] . Current Biology, 2009, 19(13):1102-1105. [49] Shine R, Warner DA, Radder R.Windows of embryonic sexual lability in two lizard species with environmental sex determination[J] . Ecology, 2007, 88(7):1781-1788. [50] Bachtrog D, Mank JE, Peichel CL, Kirkpatrick M.Sex determination:why so many ways of doing it?[J] . PLoS Biology, 2014, 12(7):1001899. [51] Organ CL, Janes DE.Evolution of sex chromosomes in Sauropsida[J] . Integrative and Comparative Biology, 2008, 48(4):512-519. [52] Graves JA.Sex chromosome specialization and degeneration in mammals[J] . Cell, 2006, 124(5):901-914. [53] Sabath N, Itescu Y, Feldman A, et al.Sex determination, longevity, and the birth and death of reptilian species[J] . Ecology and Evolution, 2016, 6(15):5207-5220. [54] Sarre SD, Ezaz T, Georges A.Transitions between sex-determining systems in reptiles and amphibians[J] . Annual review of genomics and human genetics, 2011, 12:391-406. [55] Pokorna M, Kratochvil L.Phylogeny of sex-determining mechanisms in squamate reptiles:are sex chromosomes an evolutionary trap[J] ? Zoological Journal of The Linnean Society, 2009, 156(1):168-183. [56] Valenzuela N, Adams DC.Chromosome number and sex determination coevolve in turtles[J] . Evolution, 2011, 65(6):1808-1813. [57] Badenhorst D, Hillier LW, Literman R, et al.Physical Mapping and Refinement of the Painted Turtle Genome(Chrysemys picta)Inform Amniote Genome Evolution and Challenge Turtle-Bird Chromosomal Conservation[J] . Genome Biology and Evolution, 2015, 7(7):2038-2050. [58] Janes DE, Organ C, Valenzuela N.New resources inform study of genome size, content, and organization in nonavian reptiles[J] . Integrative and Comparative Biology, 2008, 48(4):447-453. [59] Radhakrishnan S, Literman R, Neuwald J, et al.Transcriptomic responses to environmental temperature by turtles with temperature-dependent and genotypic sex determination assessed by RNAseq inform the genetic architecture of embryonic gonadal development[J] . PLoS One, 2017, 12(3):e0172044. [60] Uno Y, Nishida C, Yoshimoto S, et al.Diversity in the origins of sex chromosomes in anurans inferred from comparative mapping of sexual differentiation genes for three species of the Raninae and Xenopodinae[J] . Chromosome Research, 2008, 16(7):999-1011. [61] Matsuba C, Miura I, Merilä J.Disentangling genetic vs. environmental causes of sex determination in the common frog, Rana temporaria[J] . BMC Genetics, 2008. [62] Janes DE, Wayne ML.Evidence for a genotype x environment interaction in sex-determining response to incubation temperature in the leopard gecko, Eublepharis macularius[J] . Herpetologica, 2006, 62(1):56-62. [63] Quinn AE, Georges A, Sarre SD, et al.Temperature sex reversal implies sex gene dosage in a reptile[J] . Science, 2007, 316(5823):411-411. [64] Ospina-Alvarez N, Piferrer F.Temperature-dependent sex determination in fish revisited:prevalence, a single sex ratio response pattern, and possible effects of climate change[J] . PLoS One, 2008, 3(7):e2837. [65] Bergero R, Charlesworth D.The evolution of restricted recombination in sex chromosomes[J] . Trends in Ecology & Evolution, 2009, 24(2):94-102. [66] Miura I, Ohtani H, Nakamura M, et al.The origin and differentiation of the heteromorphic sex chromosomes Z, W, X, and Y in the frog Rana rugosa, inferred from the sequences of a sex-linked gene, ADP/ATP translocase[J] . Molecular Biology and Evolution, 1998, 15(12):1612-1619. [67] Marshall Graves JA.Weird animal genomes and the evolution of vertebrate sex and sex chromosomes[J] . Annual Review of Genetics, 2008, 42:565-586. [68] Wang C, Tang X, Xin Y, et al.Identification of Sex Chromosomes by Means of Comparative Genomic Hybridization in a Lizard, Eremias multiocellata[J] . Zoological Science, 2015, 32(2):151-156. [69] Mork L, Czerwinski M, Capel B.Predetermination of sexual fate in a turtle with temperature-dependent sex determination[J] . Developmental Biology, 2014, 386(1):264-271. [70] Radder RS, Quinn AE, Georges A, et al.Genetic evidence for co-occurrence of chromosomal and thermal sex-determining systems in a lizard[J] . Biology Letters, 2008, 4(2):176-178. [71] Barske LA, Capel B.Blurring the edges in vertebrate sex determination[J] . Current Opinion in Genetics & Development, 2008, 18(6):499-505. [72] Bachtrog D, Kirkpatrick M, Mank JE, et al.Are all sex chromosomes created equal[J] ? Trends in Genetics, 2011, 27(9):350-357. [73] Martinez PA, Ezaz T, Valenzuela N, et al.An XX/XY heteromorphic sex chromosome system in the Australian chelid turtle Emydura macquarii:a new piece in the puzzle of sex chromosome evolution in turtles[J] . Chromosome Research, 2008, 16(6):815-825. [74] Tanaka SS, Nishinakamura R.Regulation of male sex determina-tion:genital ridge formation and Sry activation in mice[J] . Cellular and Molecular Life Sciences, 2014, 71(24):4781-4802. [75] Shoemaker CM, Queen J, Crews D.Response of candidate sex-determining genes to changes in temperature reveals their involvement in the molecular network underlying temperature-dependent sex determination[J] . Molecular Endocrinology, 2007, 21(11):2750-2763. [76] Eggers S, Ohnesorg T, Sinclair A.Genetic regulation of mammalian gonad development[J] . Nature Reviews Endocrinology, 2014, 10(11):673-683. [77] Sekido R, Lovell-Badge R.Sex determination involves synergistic action of SRY and SF1 on a specific Sox9 enhancer[J] . Nature, 2008, 453(7197):930-934. [78] Pallotta MM, Turano M, Ronca R, et al.Brain Gene Expression is Influenced by Incubation Temperature During Leopard Gecko(Eublepharis macularius)Development[J] . Journal of Experimental Zoology Part B- Molecular and Developmental Evolution, 2017, 328(4):360-370. [79] Matson CK, Murphy MW, Sarver AL, et al.DMRT1 prevents female reprogramming in the postnatal mammalian testis[J] . Nature, 2011, 476(7358):101-114. [80] 包海声. Cyp19a1和Foxl2基因在中华鳖早期卵巢分化中的功能研究[D]. 上海:上海海洋大学, 2017. [81] 蔡晗, 孙伟, 刘豪彬, 包海声, 张海艳, 钱国英, 葛楚天. 中华鳖Amh基因的克隆、表达及在雄性分化中的功能初[J]. 中国科学:生命科学, 2016, 46(12):1423-1433. [82] Sun W, Cai H, Zhang G, et al.Dmrt1 is required for primary male sexual differentiation in Chinese soft-shelled turtle Pelodiscus sinensis[J] . Scientific Reports, 2017, 7(1):4433. [83] Díaz-Hernández V, Marmolejo-Valencia A, Merchant-Larios H.Exogenous estradiol alters gonadal growth and timing of temperature sex determination in gonads of sea turtle[J] . Developmental Biology, 2015, 408(1):79-89. [84] Mitchell NJ, Janzen FJ.Temperature-dependent sex determination and contemporary climate change[J] . Sexual Development, 2010, 4(1-2):129-140. [85] Telemeco RS, Elphick MJ, Shine R.Nesting lizards(Bassiana duperreyi)compensate partly, but not completely, for climate change[J] . Ecology, 2009, 90(1):17-22. |